Morphological Features

Body surface (Fig. 1)

Generic characters. With very fine reticulate sculpture in antennae, head and mesosoma (Lytoxysta, Fig. 1c) / smooth all other genera (Figs 1a, b) (except some Phaenoglyphis species, which have some very fine imbricate sulpture in scutum).


Head (Fig. 2)

Specific characters. Radial carinae on face (only for Apocharips hansoni, Fig. 2b) / smooth (Fig. 2a).


Antenna (Fig. 3)

Generic characters. Number of flagellomeres in female-male: 9-10 (Thoreauana, Fig. 3h) / 10-11 (Dilapothor, Fig. 3j) / 11-11 (Lytoxysta, Fig. 3f) / 11-12 (all other). Shape of last two flagellomeres: wider than the rest and broadly jointed (Apocharips, Dilapothor, Dilyta, Thoreauana, Figs 3g, h, i, j) / wide as the previous with constriction between them (Alloxysta, Lobopterocharips, Lytoxysta, Phaenoglyphis, Figs 3a, b, c, d, e, f). Pedicel: cup-shaped (Lobopterocharips, Fig. 3e) / cylindrical (all other).

Specific characters. Proportions (length and width) of pedicel, F1, F2, F3 and F4. Number of flagellomeres forming of club (in some species some flagellomeres are wider resembling a club). Number of flagellomeres with rhinaria. Males: F1, F2, F3 modified or not (curved, excavated, humped) (Figs 3b, d).


Pronotum (Fig. 4)

Specific characters. Lateral carinae: absent (Fig. 4c) or present (short (Fig. 4b) or long (Fig. 4a), reaching mesoscutum or not).


Mesopleura (Fig. 1)

Generic characters. Mesopleural triangle absent (Lytoxysta, Fig. 1c) / present (all other). Mesopleural sulcus present (Phaenoglyphis, Fig. 1b) / absent (all other).

Mesoscutum (Fig. 5)

Generic characters. Notauli present (only in †Protocharips and some Phaenoglyphis, Fig. 5a) / absent (all other).

Specific characters. Notauli present (Fig. 5a) or absent (Fig. 5d) (only for Phaenoglyphis species). Presence or absence of very fine imbricate sculpture in basal areas of scutum (for Phaenoglyphis species, Fig. 5c).


Scutellum (Figs 6, 7)

Generic characters. Scutellar foveae present (only in some Phaenoglyphis, Fig. 6) / absent (all other). Posterodorsal extensions of axillar strip present (Alloxysta, Lobopterocharips, Lytoxysta,

Phaenoglyphis, Fig. 10a) / absent (Apocharips, Dilapothor, Dilyta, Thoreauana, Fig. 10b). Carinae on scutellum apex: absent (Phaenoglyphis, Lobopterocharips and some Alloxysta, Fig. 7c) / longitudinal carinae at centre (some Alloxysta, Fig. 7a) / irregular carinae (Lytoxysta, Fig. 7f) / M-shaped carinae at centre (Apocharips, Fig. 7i) / ∩-shaped carinae or two long symmetrical carinae (Dilyta, Figs 7d, e) / two short symmetrical carinae (Thoreauana, Fig. 7h) / three small carinae at each side (Dilapothor, Fig. 7g).

Specific characters. Scutellar foveae absent (Fig. 6c) or present (fused, Fig. 6b or not, Figs 6a, 6b) (for Phaenoglyphis). Carinae on scutellum apex: absent (Fig. 7c) or present (from a single carina, Fig. 7a to several longitudinal carinae, Fig. 7b) (for Alloxysta species); two long symmetrical carinae (African Dilyta, Fig. 7e) or ∩-shaped carinae (Dilyta of the rest of the world, Fig. 7d).


Propodeum (Fig. 7)

Specific characters. Presence or absence of longitudinal carinae; if present, shape of carinae (short, long, thin, broad, forming a plate). Pubescence.


Forewing (Fig. 8)

Generic characters. Undulation in posteroapical margin of wing present (Lobopterocharips, Fig. 8b) / absent (all other, Fig. 8a, c). Areola present (only in †Protocharips, Fig. 5a) / absent (all other).

Specific characters. Shape, size and length of radial cell (Fig. 8).


Metasoma. (Fig. 9)

Generic characters. Visible metasoma with two large terga with subequal dorsomedian lengths (Alloxysta, Lobopterocharips, Lytoxysta, Phaenoglyphis, Fig. 9a; also in †Protocharips, Fig. 5a) / with a small basal tergum, terminating just posterior to ring of setae (Apocharips, Fig. 9b) / not segmented, only one tergite visible (Dilapothor, Dilyta, Thoreauana, Figs 9c, d).

Specific characters. Puntuation on distal area absent (Fig. 9c) or present (Fig. 9d) (for Dilyta species).